DK952029
Clone id TST38A01NGRL0012_P21
Library
Length 659
Definition Adiantum capillus-veneris mRNA. clone: TST38A01NGRL0012_P21. 5' end sequence.
Accession
Tissue type prothallia
Developmental stage gametophyte
Contig ID
Sequence
AGATATGTTACCATGTCTTGGGACTGACACATCAGATGTGGTCACCAGTTCCACAGTCGA
TGAAAGATTACATTGCCACTCCAAAACCTAATGGATATCAAAGTCTTCACACAAAAGTGC
TTCCATTTCTTTATGAAAATACTTTTCGCCTAGAGATCCAGATTCGATCAGAGGATATGG
ATATTATTGCCGAGCGAGGCATTGCTGCACATTACAGTGGTAGGACTGTCTGCACATCCT
TGTCTACAGAAGCAGGTTCACACAAAGGATCCAGCAGTTTCATGAACAATGTTGATCTAG
CACGTAGGGTTAGCTGGCTTAACGCAATACGGGAATGGCAAGAGGAGTTTGTTGGAAATA
TGTCTGCCCAGGAGTTTGTCGATACTGTCACTGGGGATCTTCTTGGTAGTCGTGTATTTG
TTTTCACACCGAAAGGAGAAATTAAGAACCTTCCCAAGGGTGCAACAGTCATTGATTATG
CTTACCAAATACATACAGACTTGGGAAATAACATGGTGGCTGCAAAGGTAAATGGAAATT
TGGTCTCACCGACACATACATTGGCCAACGCTGAAGTGGTTGAAGTTATCACCTATAATG
GTTTATCAATAAAATCAGTTTTTCATCGACACAGGCAATGGTTGCAGCATGCAAAAACA
■■Homology search results ■■ -
sp_hit_id O52177
Definition sp|O52177|RELA_MYXXA GTP pyrophosphokinase OS=Myxococcus xanthus
Align length 218
Score (bit) 171.0
E-value 2.0e-42
Report
BLASTX 2.2.19 [Nov-02-2008]


Reference: Altschul, Stephen F., Thomas L. Madden, Alejandro A. Schaffer,
Jinghui Zhang, Zheng Zhang, Webb Miller, and David J. Lipman (1997),
"Gapped BLAST and PSI-BLAST: a new generation of protein database search
programs", Nucleic Acids Res. 25:3389-3402.

Query= DK952029|Adiantum capillus-veneris mRNA, clone:
TST38A01NGRL0012_P21, 5'
(659 letters)

Database: uniprot_sprot.fasta
412,525 sequences; 148,809,765 total letters

Searching..................................................done



Score E
Sequences producing significant alignments: (bits) Value

sp|O52177|RELA_MYXXA GTP pyrophosphokinase OS=Myxococcus xanthus... 171 2e-42
sp|O54408|RELA_BACSU GTP pyrophosphokinase OS=Bacillus subtilis ... 153 7e-37
sp|P74007|SPOT_SYNY3 Probable guanosine-3',5'-bis(diphosphate) 3... 153 9e-37
sp|P52560|RELA_STRCO GTP pyrophosphokinase OS=Streptomyces coeli... 147 4e-35
sp|Q54089|RELA_STREQ Putative GTP pyrophosphokinase OS=Streptoco... 146 8e-35
sp|Q8CS97|RELA_STAES GTP pyrophosphokinase OS=Staphylococcus epi... 145 2e-34
sp|Q5HNR8|RELA_STAEQ GTP pyrophosphokinase OS=Staphylococcus epi... 145 2e-34
sp|O51216|SPOT_BORBU Guanosine-3',5'-bis(diphosphate) 3'-pyropho... 144 3e-34
sp|P0A0E9|RELA_STAAW GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|P0A0F0|RELA_STAAU GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|Q6G8T5|RELA_STAAS GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|Q6GG70|RELA_STAAR GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|Q99TL8|RELA_STAAN GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|Q931Q4|RELA_STAAM GTP pyrophosphokinase OS=Staphylococcus aur... 144 4e-34
sp|O85709|RELA_STRAT GTP pyrophosphokinase OS=Streptomyces antib... 143 7e-34
sp|P0AG23|RELA_SHIFL GTP pyrophosphokinase OS=Shigella flexneri ... 137 5e-32
sp|P0AG20|RELA_ECOLI GTP pyrophosphokinase OS=Escherichia coli (... 137 5e-32
sp|P0AG21|RELA_ECOL6 GTP pyrophosphokinase OS=Escherichia coli O... 137 5e-32
sp|P0AG22|RELA_ECO57 GTP pyrophosphokinase OS=Escherichia coli O... 137 5e-32
sp|P0AG26|SPOT_SHIFL Guanosine-3',5'-bis(diphosphate) 3'-pyropho... 136 9e-32
sp|P0AG24|SPOT_ECOLI Guanosine-3',5'-bis(diphosphate) 3'-pyropho... 136 9e-32
sp|P0AG25|SPOT_ECO57 Guanosine-3',5'-bis(diphosphate) 3'-pyropho... 136 9e-32
sp|O67012|SPOT_AQUAE Guanosine-3',5'-bis(diphosphate) 3'-pyropho... 135 2e-31
sp|Q8CY42|RSH_BRUSU GTP pyrophosphokinase rsh OS=Brucella suis G... 135 2e-31
sp|A5VPI9|RSH_BRUO2 GTP pyrophosphokinase rsh OS=Brucella ovis (... 135 2e-31
sp|Q8YG65|RSH_BRUME GTP pyrophosphokinase rsh OS=Brucella melite... 135 2e-31
sp|Q57E90|RSH_BRUAB GTP pyrophosphokinase rsh OS=Brucella abortu... 135 2e-31
sp|Q2YN11|RSH_BRUA2 GTP pyrophosphokinase rsh OS=Brucella abortu... 135 2e-31
sp|P66014|RELA_MYCTU Probable GTP pyrophosphokinase OS=Mycobacte... 134 4e-31
sp|P66015|RELA_MYCBO Probable GTP pyrophosphokinase OS=Mycobacte... 134 4e-31

>sp|O52177|RELA_MYXXA GTP pyrophosphokinase OS=Myxococcus xanthus
GN=relA PE=3 SV=1
Length = 757

Score = 171 bits (434), Expect = 2e-42
Identities = 92/218 (42%), Positives = 128/218 (58%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY LGL H+MW PVP KD+IA PKPN YQSLHT ++ L E R+E+QIR+ +M
Sbjct: 294 CYEALGLVHEMWKPVPGRFKDFIAIPKPNMYQSLHTTIIGPLSE---RVEVQIRTSEMHK 350

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE GIAAH+ +K + ++ D + +WL + EWQ++
Sbjct: 351 IAEEGIAAHWK--------------YKEGKAVISKDD--EKFAWLRQLMEWQQDL---KD 391

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
+EF++TV DL VFVFTPKG++++LP+GAT +D+AY IH+D+GN V AKVNG +V
Sbjct: 392 PKEFLETVKVDLFTDEVFVFTPKGDVRSLPRGATPVDFAYAIHSDVGNRCVGAKVNGKIV 451

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ + N + VEV+T H + WL KT
Sbjct: 452 PLRYKMKNGDTVEVLT------SPQQHPSKDWLTFVKT 483


>sp|O54408|RELA_BACSU GTP pyrophosphokinase OS=Bacillus subtilis
GN=relA PE=3 SV=2
Length = 734

Score = 153 bits (387), Expect = 7e-37
Identities = 82/218 (37%), Positives = 125/218 (57%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY VLG+ H W P+P KDYIA PKPN YQSLHT V+ LE+QIR+ +M
Sbjct: 278 CYAVLGIIHTCWKPMPGRFKDYIAMPKPNMYQSLHTTVIG---PKADPLEVQIRTFEMHE 334

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE G+AAH++ +K + +++SW I E+Q E +
Sbjct: 335 IAEYGVAAHWA--------------YKEGKAANEGATFEKKLSWFREILEFQNE---STD 377

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
A+EF++++ DL V+VFTPKG++ LP G+ ID++Y+IH+++GN + AKVNG +V
Sbjct: 378 AEEFMESLKIDLFSDMVYVFTPKGDVIELPSGSVPIDFSYRIHSEIGNKTIGAKVNGKMV 437

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ H L ++VE++T K + + W++ A+T
Sbjct: 438 TLDHKLRTGDIVEILT-----SKHSYGPSQDWVKLAQT 470


>sp|P74007|SPOT_SYNY3 Probable guanosine-3',5'-bis(diphosphate)
3'-pyrophosphohydrolase OS=Synechocystis sp. (strain PCC
6803) GN=spoT PE=3 SV=1
Length = 760

Score = 153 bits (386), Expect = 9e-37
Identities = 85/227 (37%), Positives = 131/227 (57%), Gaps = 10/227 (4%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY L + H ++ P+P KDYI PKPN YQSLHT VL + LEIQIR+E+M
Sbjct: 305 CYRALSVVHDVFKPIPGRFKDYIGLPKPNRYQSLHTTVLGL---TSRPLEIQIRTEEMHH 361

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
+AE GIAAH+ + E+G + ++ + + D + +WL + +WQ +
Sbjct: 362 VAEYGIAAHWKYK--------ESGGSENAT--LTSTD--EKFTWLRQLLDWQSDL---KD 406

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
AQE+V+ + +L V+VFTPKGE+ +L +GAT +D+AY+IHT++G++M A+VNG +
Sbjct: 407 AQEYVENLKQNLFDDDVYVFTPKGEVISLARGATPVDFAYRIHTEVGHHMKGARVNGQWL 466

Query: 546 SPTHTLANAEVVEVIT----------YNGLSIKSVFHRHRQWLQHAK 656
L N ++VE++T N + S HR RQW + ++
Sbjct: 467 GVDTRLKNGDIVEIVTQKNSHPSLDWLNFVVTPSARHRIRQWFKRSR 513


>sp|P52560|RELA_STRCO GTP pyrophosphokinase OS=Streptomyces
coelicolor GN=relA PE=2 SV=1
Length = 847

Score = 147 bits (372), Expect = 4e-35
Identities = 81/196 (41%), Positives = 116/196 (59%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY LG H W+PVP KDYIA PK N YQSLHT V+ +E+QIR+ DM
Sbjct: 375 CYAALGTVHARWNPVPGRFKDYIAMPKFNMYQSLHTTVIG---PGGKPVELQIRTFDMHR 431

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
AE GIAAH+ + + +++ + SS + D ++WL + +WQ+E
Sbjct: 432 RAEYGIAAHWKYKQEAVAGASKVRTDAPKSSGKSKDDHLNDMAWLRQLLDWQKE---TED 488

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
EF++++ DL + VFVFTPKG++ LP GAT +D+AY +HT++G+ + A+VNG LV
Sbjct: 489 PGEFLESLRFDLSRNEVFVFTPKGDVIALPAGATPVDFAYAVHTEVGHRTIGARVNGRLV 548

Query: 546 SPTHTLANAEVVEVIT 593
TL N ++VEV T
Sbjct: 549 PLESTLDNGDLVEVFT 564


>sp|Q54089|RELA_STREQ Putative GTP pyrophosphokinase
OS=Streptococcus equisimilis GN=relA PE=1 SV=1
Length = 739

Score = 146 bits (369), Expect = 8e-35
Identities = 81/217 (37%), Positives = 121/217 (55%)
Frame = +3

Query: 9 YHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDII 188
Y ++G H++W P+P KDYIA PK NGYQS+HT V Y +EIQIR+++M +
Sbjct: 279 YAMVGYIHELWRPMPGRFKDYIAAPKANGYQSIHTTV----YGPKGPIEIQIRTKEMHQV 334

Query: 189 AERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMSA 368
AE G+AAH++ + KG +N + ++W+ + E Q+ N A
Sbjct: 335 AEYGVAAHWAYK-------------KGVRGKVNQAEQKVGMNWIKELVELQD--ASNGDA 379

Query: 369 QEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLVS 548
+FVD+V D+ R++VFTP G ++ LPK + ID+AY IHT +G + AKVNG +V
Sbjct: 380 VDFVDSVKEDIFSERIYVFTPTGAVQELPKDSGPIDFAYAIHTQVGEKAIGAKVNGRMVP 439

Query: 549 PTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
T L +VVE++T + F R W++ KT
Sbjct: 440 LTAKLKTGDVVEIVTN-----PNSFGPSRDWIKLVKT 471


>sp|Q8CS97|RELA_STAES GTP pyrophosphokinase OS=Staphylococcus
epidermidis (strain ATCC 12228) GN=relA PE=3 SV=1
Length = 729

Score = 145 bits (366), Expect = 2e-34
Identities = 81/218 (37%), Positives = 119/218 (54%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY LGL H +W P+P KDYIA PK N YQSLHT V+ N LEIQIR+ +M
Sbjct: 278 CYATLGLVHTLWKPMPGRFKDYIAMPKQNMYQSLHTTVVG---PNGDPLEIQIRTHEMHE 334

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE G+AAH++ + +G + D +++WL + E +
Sbjct: 335 IAEHGVAAHWAYK-------------EGKTVNQKTQDFQNKLNWLKELAETDHT---SAD 378

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
AQEF++++ DL +V+ FTP ++ LP GA ID+AY IH+++GN M+ AKVNG +V
Sbjct: 379 AQEFMESLKYDLQSDKVYAFTPASDVIELPYGAVPIDFAYAIHSEVGNKMIGAKVNGKIV 438

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ L +++E+ T K + R WL+ K+
Sbjct: 439 PIDYVLQTGDIIEIRT-----SKHSYGPSRDWLKIVKS 471


>sp|Q5HNR8|RELA_STAEQ GTP pyrophosphokinase OS=Staphylococcus
epidermidis (strain ATCC 35984 / RP62A) GN=relA PE=3
SV=1
Length = 729

Score = 145 bits (366), Expect = 2e-34
Identities = 81/218 (37%), Positives = 119/218 (54%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY LGL H +W P+P KDYIA PK N YQSLHT V+ N LEIQIR+ +M
Sbjct: 278 CYATLGLVHTLWKPMPGRFKDYIAMPKQNMYQSLHTTVVG---PNGDPLEIQIRTHEMHE 334

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE G+AAH++ + +G + D +++WL + E +
Sbjct: 335 IAEHGVAAHWAYK-------------EGKTVNQKTQDFQNKLNWLKELAETDHT---SAD 378

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
AQEF++++ DL +V+ FTP ++ LP GA ID+AY IH+++GN M+ AKVNG +V
Sbjct: 379 AQEFMESLKYDLQSDKVYAFTPASDVIELPYGAVPIDFAYAIHSEVGNKMIGAKVNGKIV 438

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ L +++E+ T K + R WL+ K+
Sbjct: 439 PIDYVLQTGDIIEIRT-----SKHSYGPSRDWLKIVKS 471


>sp|O51216|SPOT_BORBU Guanosine-3',5'-bis(diphosphate)
3'-pyrophosphohydrolase OS=Borrelia burgdorferi GN=spoT
PE=3 SV=1
Length = 667

Score = 144 bits (364), Expect = 3e-34
Identities = 81/224 (36%), Positives = 127/224 (56%), Gaps = 11/224 (4%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY +L + H++W P+P +KDYIA+PK N YQSLHT V + E+ +EIQIR+E+MD
Sbjct: 290 CYEILEIVHRVWKPIPGRLKDYIASPKENKYQSLHTTVR--IPEDNQLIEIQIRTEEMDR 347

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDL-ARRVSWLNAIREWQEEFVGNM 362
IA G+AAH+ + ++L A +S++N I++WQ+E
Sbjct: 348 IANYGVAAHWI--------------------YKEQIELKADDLSFINRIKKWQQESANK- 386

Query: 363 SAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNL 542
+Q ++ + +LL + ++V+TP+GE+ LP G+ ID+AY IHTD+G+ + AK+NG +
Sbjct: 387 -SQYSMNDIHKELLNTFIYVYTPEGEVVELPFGSNSIDFAYIIHTDIGDQALYAKINGKI 445

Query: 543 VSPTHTLANAEVVEVIT----------YNGLSIKSVFHRHRQWL 644
S T L N ++VE+ T N + K + R WL
Sbjct: 446 SSITKPLKNEQIVEIFTSKDSKPDVIWLNSVRTKKARSKIRSWL 489


>sp|P0A0E9|RELA_STAAW GTP pyrophosphokinase OS=Staphylococcus aureus
(strain MW2) GN=relA PE=3 SV=1
Length = 736

Score = 144 bits (363), Expect = 4e-34
Identities = 83/218 (38%), Positives = 119/218 (54%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY +LGL H +W P+P KDYIA PK N YQSLHT V+ N LEIQIR+ DM
Sbjct: 285 CYAILGLVHTLWKPMPGRFKDYIAMPKQNLYQSLHTTVVG---PNGDPLEIQIRTFDMHE 341

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE G+AAH++ + +G + +++WL + E +
Sbjct: 342 IAEHGVAAHWAYK-------------EGKKVSEKDQTYQNKLNWLKELAEADHT---SSD 385

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
AQEF++T+ DL +V+ FTP ++ LP GA ID+AY IH+++GN M+ AKVNG +V
Sbjct: 386 AQEFMETLKYDLQSDKVYAFTPASDVIELPYGAVPIDFAYAIHSEVGNKMIGAKVNGKIV 445

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ L ++VE+ T K + R WL+ K+
Sbjct: 446 PIDYILQTGDIVEIRT-----SKHSYGPSRDWLKIVKS 478


>sp|P0A0F0|RELA_STAAU GTP pyrophosphokinase OS=Staphylococcus aureus
GN=relA PE=3 SV=1
Length = 736

Score = 144 bits (363), Expect = 4e-34
Identities = 83/218 (38%), Positives = 119/218 (54%)
Frame = +3

Query: 6 CYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDI 185
CY +LGL H +W P+P KDYIA PK N YQSLHT V+ N LEIQIR+ DM
Sbjct: 285 CYAILGLVHTLWKPMPGRFKDYIAMPKQNLYQSLHTTVVG---PNGDPLEIQIRTFDMHE 341

Query: 186 IAERGIAAHYSGRTVCTSLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMS 365
IAE G+AAH++ + +G + +++WL + E +
Sbjct: 342 IAEHGVAAHWAYK-------------EGKKVSEKDQTYQNKLNWLKELAEADHT---SSD 385

Query: 366 AQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLV 545
AQEF++T+ DL +V+ FTP ++ LP GA ID+AY IH+++GN M+ AKVNG +V
Sbjct: 386 AQEFMETLKYDLQSDKVYAFTPASDVIELPYGAVPIDFAYAIHSEVGNKMIGAKVNGKIV 445

Query: 546 SPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
+ L ++VE+ T K + R WL+ K+
Sbjct: 446 PIDYILQTGDIVEIRT-----SKHSYGPSRDWLKIVKS 478


tr_hit_id B8LK81
Definition tr|B8LK81|B8LK81_PICSI Putative uncharacterized protein OS=Picea sitchensis
Align length 220
Score (bit) 349.0
E-value 8.0e-95
Report
BLASTX 2.2.19 [Nov-02-2008]


Reference: Altschul, Stephen F., Thomas L. Madden, Alejandro A. Schaffer,
Jinghui Zhang, Zheng Zhang, Webb Miller, and David J. Lipman (1997),
"Gapped BLAST and PSI-BLAST: a new generation of protein database search
programs", Nucleic Acids Res. 25:3389-3402.

Query= DK952029|Adiantum capillus-veneris mRNA, clone:
TST38A01NGRL0012_P21, 5'
(659 letters)

Database: uniprot_trembl.fasta
7,341,751 sequences; 2,391,615,440 total letters

Searching..................................................done



Score E
Sequences producing significant alignments: (bits) Value

tr|B8LK81|B8LK81_PICSI Putative uncharacterized protein OS=Picea... 349 8e-95
tr|Q9M5P7|Q9M5P7_ARATH RSH1 OS=Arabidopsis thaliana GN=At4g02260... 345 1e-93
tr|Q3EAC8|Q3EAC8_ARATH Uncharacterized protein At4g02260.3 OS=Ar... 345 1e-93
tr|Q84LE7|Q84LE7_TOBAC RelA-SpoT like protein RSH1 OS=Nicotiana ... 343 7e-93
tr|Q0WUY0|Q0WUY0_ARATH RSH1 OS=Arabidopsis thaliana GN=At4g02260... 342 9e-93
tr|Q9AYT5|Q9AYT5_ORYSJ Os03g0340900 protein OS=Oryza sativa subs... 331 3e-89
tr|A9TAM9|A9TAM9_PHYPA Predicted protein (Fragment) OS=Physcomit... 327 4e-88
tr|A9TAK9|A9TAK9_PHYPA Predicted protein (Fragment) OS=Physcomit... 321 2e-86
tr|Q9M112|Q9M112_ARATH Putative GTP pyrophosphokinase (Fragment)... 316 7e-85
tr|O81418|O81418_ARATH T2H3.9 OS=Arabidopsis thaliana GN=T2H3.9 ... 311 3e-83
tr|A7Q0G3|A7Q0G3_VITVI Chromosome chr7 scaffold_42, whole genome... 297 3e-79
tr|B8APD8|B8APD8_ORYSI Putative uncharacterized protein OS=Oryza... 280 7e-74
tr|A3AHS8|A3AHS8_ORYSJ Putative uncharacterized protein OS=Oryza... 258 2e-67
tr|Q72HX7|Q72HX7_THET2 Guanosine-3',5'-bis(Diphosphate) 3'-pyrop... 206 1e-51
tr|Q5SHL3|Q5SHL3_THET8 Guanosine-3',5'-bis(Diphosphate) 3'-pyrop... 206 1e-51
tr|B7A923|B7A923_THEAQ (P)ppGpp synthetase I, SpoT/RelA OS=Therm... 203 7e-51
tr|A4RY78|A4RY78_OSTLU Predicted protein OS=Ostreococcus lucimar... 177 4e-43
tr|Q3ZAI9|Q3ZAI9_DEHE1 GTP pyrophosphokinase OS=Dehalococcoides ... 177 5e-43
tr|A8CST3|A8CST3_9CHLR (P)ppGpp synthetase I, SpoT/RelA OS=Dehal... 177 5e-43
tr|Q3ZW91|Q3ZW91_DEHSC GTP pyrophosphokinase OS=Dehalococcoides ... 177 7e-43
tr|A5FSR6|A5FSR6_DEHSB (P)ppGpp synthetase I, SpoT/RelA OS=Dehal... 177 7e-43
tr|Q096V2|Q096V2_STIAU RelA/SpoT family protein OS=Stigmatella a... 174 6e-42
tr|Q1D7G8|Q1D7G8_MYXXD GTP pyrophosphokinase OS=Myxococcus xanth... 171 3e-41
tr|A7H9X3|A7H9X3_ANADF (P)ppGpp synthetase I, SpoT/RelA OS=Anaer... 167 4e-40
tr|Q3A9Z8|Q3A9Z8_CARHZ GTP pyrophosphokinase OS=Carboxydothermus... 167 7e-40
tr|Q2IKY6|Q2IKY6_ANADE (P)ppGpp synthetase I (GTP pyrophosphokin... 164 3e-39
tr|B4UIC6|B4UIC6_ANASK (P)ppGpp synthetase I, SpoT/RelA OS=Anaer... 164 3e-39
tr|B8JH42|B8JH42_9DELT (P)ppGpp synthetase I, SpoT/RelA OS=Anaer... 164 3e-39
tr|B7DNW0|B7DNW0_9BACL (P)ppGpp synthetase I, SpoT/RelA OS=Alicy... 164 3e-39
tr|B3E4R4|B3E4R4_GEOLS (P)ppGpp synthetase I, SpoT/RelA OS=Geoba... 164 4e-39

>tr|B8LK81|B8LK81_PICSI Putative uncharacterized protein OS=Picea
sitchensis PE=2 SV=1
Length = 901

Score = 349 bits (896), Expect = 8e-95
Identities = 162/220 (73%), Positives = 196/220 (89%), Gaps = 1/220 (0%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYH+LGL H +W+P+P+ MKDYIATPKPNGYQSL TKV+PFL E+TFR+E+QIR+E+MD
Sbjct: 432 ICYHILGLVHGIWTPIPREMKDYIATPKPNGYQSLQTKVIPFLNESTFRVEVQIRTEEMD 491

Query: 183 IIAERGIAAHYSGRTVCT-SLSTEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGN 359
IA+RGIA HYSG+ + + S T +G ++ +N+ D+ARRVSWLNAIREWQEEFVGN
Sbjct: 492 CIAQRGIAVHYSGKGITSGSARTNGRILRGKATCLNSADIARRVSWLNAIREWQEEFVGN 551

Query: 360 MSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGN 539
MS++EFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATV+DYAYQ+HT++GN M+AAKVNGN
Sbjct: 552 MSSREFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVVDYAYQVHTEIGNKMIAAKVNGN 611

Query: 540 LVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
VSP HTLANAEVVE+ITY+GLS K+VFHRH+QWLQ+A+T
Sbjct: 612 PVSPMHTLANAEVVEIITYDGLSSKTVFHRHQQWLQYART 651


>tr|Q9M5P7|Q9M5P7_ARATH RSH1 OS=Arabidopsis thaliana GN=At4g02260 PE=2
SV=1
Length = 883

Score = 345 bits (885), Expect = 1e-93
Identities = 161/223 (72%), Positives = 191/223 (85%), Gaps = 4/223 (1%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H++W P+P+++KDYIATPKPNGYQSLHT V+PFLYE+ FRLE+QIR+E+MD
Sbjct: 425 ICYHVLGLVHEIWKPIPRTVKDYIATPKPNGYQSLHTTVIPFLYESMFRLEVQIRTEEMD 484

Query: 183 IIAERGIAAHYSGRTVCTSLSTEA----GSHKGSSSFMNNVDLARRVSWLNAIREWQEEF 350
+IAERGIA +Y+G+++ T L A + +G + +NN D A RV WLNAIREWQEEF
Sbjct: 485 LIAERGIAVYYNGKSLSTGLVGNAVPLGRNSRGKTGCLNNADFALRVGWLNAIREWQEEF 544

Query: 351 VGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKV 530
VGNMS++EFVDT+T DLLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAKV
Sbjct: 545 VGNMSSREFVDTITRDLLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAKV 604

Query: 531 NGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
NGNLVSPTH L NAEVVE++TYN LS KS F RH+QWLQHAKT
Sbjct: 605 NGNLVSPTHVLENAEVVEIVTYNALSSKSAFQRHKQWLQHAKT 647


>tr|Q3EAC8|Q3EAC8_ARATH Uncharacterized protein At4g02260.3
OS=Arabidopsis thaliana GN=At4g02260 PE=4 SV=1
Length = 816

Score = 345 bits (885), Expect = 1e-93
Identities = 161/223 (72%), Positives = 191/223 (85%), Gaps = 4/223 (1%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H++W P+P+++KDYIATPKPNGYQSLHT V+PFLYE+ FRLE+QIR+E+MD
Sbjct: 425 ICYHVLGLVHEIWKPIPRTVKDYIATPKPNGYQSLHTTVIPFLYESMFRLEVQIRTEEMD 484

Query: 183 IIAERGIAAHYSGRTVCTSLSTEA----GSHKGSSSFMNNVDLARRVSWLNAIREWQEEF 350
+IAERGIA +Y+G+++ T L A + +G + +NN D A RV WLNAIREWQEEF
Sbjct: 485 LIAERGIAVYYNGKSLSTGLVGNAVPLGRNSRGKTGCLNNADFALRVGWLNAIREWQEEF 544

Query: 351 VGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKV 530
VGNMS++EFVDT+T DLLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAKV
Sbjct: 545 VGNMSSREFVDTITRDLLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAKV 604

Query: 531 NGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
NGNLVSPTH L NAEVVE++TYN LS KS F RH+QWLQHAKT
Sbjct: 605 NGNLVSPTHVLENAEVVEIVTYNALSSKSAFQRHKQWLQHAKT 647


>tr|Q84LE7|Q84LE7_TOBAC RelA-SpoT like protein RSH1 OS=Nicotiana
tabacum GN=NtRSH1 PE=2 SV=1
Length = 876

Score = 343 bits (879), Expect = 7e-93
Identities = 162/223 (72%), Positives = 189/223 (84%), Gaps = 4/223 (1%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H +W+P+P++MKDY+ATPKPNGYQSLHT V+PFLYE+ FRLE+QIR+E+MD
Sbjct: 418 ICYHVLGLVHGIWTPIPRAMKDYVATPKPNGYQSLHTTVIPFLYESMFRLEVQIRTEEMD 477

Query: 183 IIAERGIAAHYSGRTVCTSLS----TEAGSHKGSSSFMNNVDLARRVSWLNAIREWQEEF 350
+IAERGIAAHYSG+ L T S +G +NN ++A R+ WLNAIREWQEEF
Sbjct: 478 LIAERGIAAHYSGKGFVNGLVGHVITNGRSSRGKIVCLNNANIALRIGWLNAIREWQEEF 537

Query: 351 VGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKV 530
VGNMS++EFVDT+T DLLGSRVFVFTP GEIK+LPKGATVIDYAY IHT++GN MVAAKV
Sbjct: 538 VGNMSSREFVDTITRDLLGSRVFVFTPGGEIKHLPKGATVIDYAYMIHTEIGNKMVAAKV 597

Query: 531 NGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
NGNLVSP H LANAEVVE+ITYNGLS KS F RH++WLQHAKT
Sbjct: 598 NGNLVSPLHVLANAEVVEIITYNGLSSKSAFERHKEWLQHAKT 640


>tr|Q0WUY0|Q0WUY0_ARATH RSH1 OS=Arabidopsis thaliana GN=At4g02260 PE=2
SV=1
Length = 884

Score = 342 bits (878), Expect = 9e-93
Identities = 160/223 (71%), Positives = 190/223 (85%), Gaps = 4/223 (1%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H++W P+P+++KDYIATPKPNGYQSLHT V+PFLYE+ FRLE+QIR+E+MD
Sbjct: 426 ICYHVLGLVHEIWKPIPRTVKDYIATPKPNGYQSLHTTVIPFLYESMFRLEVQIRTEEMD 485

Query: 183 IIAERGIAAHYSGRTVCTSLSTEA----GSHKGSSSFMNNVDLARRVSWLNAIREWQEEF 350
+IAER IA +Y+G+++ T L A + +G + +NN D A RV WLNAIREWQEEF
Sbjct: 486 LIAERDIAVYYNGKSLSTGLVGNAVPLGRNSRGKTGCLNNADFALRVGWLNAIREWQEEF 545

Query: 351 VGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKV 530
VGNMS++EFVDT+T DLLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAKV
Sbjct: 546 VGNMSSREFVDTITRDLLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAKV 605

Query: 531 NGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
NGNLVSPTH L NAEVVE++TYN LS KS F RH+QWLQHAKT
Sbjct: 606 NGNLVSPTHVLENAEVVEIVTYNALSSKSAFQRHKQWLQHAKT 648


>tr|Q9AYT5|Q9AYT5_ORYSJ Os03g0340900 protein OS=Oryza sativa subsp.
japonica GN=Os-RelA1 PE=2 SV=1
Length = 892

Score = 331 bits (848), Expect = 3e-89
Identities = 161/224 (71%), Positives = 184/224 (82%), Gaps = 5/224 (2%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H +W+P+PQ++KDYIATPKPNGYQSLHT V+PFL E+ F LE+QIR+EDMD
Sbjct: 437 ICYHVLGLVHGIWTPIPQAVKDYIATPKPNGYQSLHTTVIPFLNESMFHLEVQIRTEDMD 496

Query: 183 IIAERGIAAHYSGRTVCTSLSTEAGSHKGSSS-----FMNNVDLARRVSWLNAIREWQEE 347
+IAERGIAAHYSGR V + G G +S +NN A R+ WLNAIREWQEE
Sbjct: 497 LIAERGIAAHYSGRGVVSG-PVRPGISSGRNSNGKVICLNNTGFALRIGWLNAIREWQEE 555

Query: 348 FVGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAK 527
FVGNMS++EFVDT+T DLLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAK
Sbjct: 556 FVGNMSSREFVDTITRDLLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAK 615

Query: 528 VNGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
VNGNLVSP H LANAEVVE+I Y+ LS K F RH+QWLQHAKT
Sbjct: 616 VNGNLVSPIHVLANAEVVEIIIYDKLSAKYAFQRHQQWLQHAKT 659


>tr|A9TAM9|A9TAM9_PHYPA Predicted protein (Fragment) OS=Physcomitrella
patens subsp. patens GN=PHYPADRAFT_142688 PE=4 SV=1
Length = 873

Score = 327 bits (838), Expect = 4e-88
Identities = 161/236 (68%), Positives = 187/236 (79%), Gaps = 17/236 (7%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
+CYHVLGL H MW P+PQSMKDYIATPKPNGYQSLHT V+PF + F LEIQIR+++MD
Sbjct: 388 VCYHVLGLVHAMWPPLPQSMKDYIATPKPNGYQSLHTTVIPFGTQTYFPLEIQIRTDEMD 447

Query: 183 IIAERGIAAHYSGRTVCTSLSTE-----------------AGSHKGSSSFMNNVDLARRV 311
+AE GIAA+YSGR + S A + GSSS++N+ D+ARRV
Sbjct: 448 RLAECGIAAYYSGRWMAERNSDAMVPFARNEEDAARAIYPASNSLGSSSYLNDADIARRV 507

Query: 312 SWLNAIREWQEEFVGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQI 491
SWL +IR+WQEEFVGNMS++EFVDTVTGDLL SRVFVFTPKGE+KNLPKGATVIDYAYQI
Sbjct: 508 SWLKSIRDWQEEFVGNMSSREFVDTVTGDLLSSRVFVFTPKGEVKNLPKGATVIDYAYQI 567

Query: 492 HTDLGNNMVAAKVNGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
HTD+GN MVAAKVNGNLVSPTH LANAEVVE++TY+ +S K VF H QWLQ+AKT
Sbjct: 568 HTDVGNKMVAAKVNGNLVSPTHALANAEVVEILTYDSISAKRVFQLHSQWLQYAKT 623


>tr|A9TAK9|A9TAK9_PHYPA Predicted protein (Fragment) OS=Physcomitrella
patens subsp. patens GN=PHYPADRAFT_142546 PE=4 SV=1
Length = 886

Score = 321 bits (823), Expect = 2e-86
Identities = 159/236 (67%), Positives = 187/236 (79%), Gaps = 17/236 (7%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
+CYHVLGL H MW P+PQ+MKDYIATPKPNGYQSLHT V+PF + F +EIQIR+E+MD
Sbjct: 406 VCYHVLGLVHAMWPPLPQTMKDYIATPKPNGYQSLHTTVIPFGTQTYFPIEIQIRTEEMD 465

Query: 183 IIAERGIAAHYSGRTVCTSLSTEAG--SHKG---------------SSSFMNNVDLARRV 311
+AE GIAAH+SGR V S G + KG S++ +N+ D+ARRV
Sbjct: 466 KLAEWGIAAHHSGRWVAERESDAMGPFARKGTNGAAAGNLASNPPSSNTCLNDADIARRV 525

Query: 312 SWLNAIREWQEEFVGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQI 491
SWL +IR+WQEEFVGNMS++EFVDTVTGDLL SRVFVFTPKGE+KNLPKGATVIDYAYQI
Sbjct: 526 SWLKSIRDWQEEFVGNMSSREFVDTVTGDLLSSRVFVFTPKGEVKNLPKGATVIDYAYQI 585

Query: 492 HTDLGNNMVAAKVNGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
HTD+GN MVAAKVNG LVSPTHTLANAEVVE++TY+G+S K +F HRQWL +A T
Sbjct: 586 HTDVGNKMVAAKVNGILVSPTHTLANAEVVEIVTYDGVSTKKIFQLHRQWLAYAMT 641


>tr|Q9M112|Q9M112_ARATH Putative GTP pyrophosphokinase (Fragment)
OS=Arabidopsis thaliana GN=AT4g02260 PE=2 SV=1
Length = 816

Score = 316 bits (810), Expect = 7e-85
Identities = 151/207 (72%), Positives = 176/207 (85%), Gaps = 4/207 (1%)
Frame = +3

Query: 51 PQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMDIIAERGIAAHYSGRTV 230
PQ +KDYIATPKPNGYQSLHT V+PFLYE+ FRLE+QIR+E+MD+IAERGIA +Y+G+++
Sbjct: 383 PQQVKDYIATPKPNGYQSLHTTVIPFLYESMFRLEVQIRTEEMDLIAERGIAVYYNGKSL 442

Query: 231 CTSLSTEA----GSHKGSSSFMNNVDLARRVSWLNAIREWQEEFVGNMSAQEFVDTVTGD 398
T L A + +G + +NN D A RV WLNAIREWQEEFVGNMS++EFVDT+T D
Sbjct: 443 STGLVGNAVPLGRNSRGKTGCLNNADFALRVGWLNAIREWQEEFVGNMSSREFVDTITRD 502

Query: 399 LLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKVNGNLVSPTHTLANAEV 578
LLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAKVNGNLVSPTH L NAEV
Sbjct: 503 LLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAKVNGNLVSPTHVLENAEV 562

Query: 579 VEVITYNGLSIKSVFHRHRQWLQHAKT 659
VE++TYN LS KS F RH+QWLQHAKT
Sbjct: 563 VEIVTYNALSSKSAFQRHKQWLQHAKT 589


>tr|O81418|O81418_ARATH T2H3.9 OS=Arabidopsis thaliana GN=T2H3.9
PE=4 SV=1
Length = 615

Score = 311 bits (796), Expect = 3e-83
Identities = 150/223 (67%), Positives = 177/223 (79%), Gaps = 4/223 (1%)
Frame = +3

Query: 3 ICYHVLGLTHQMWSPVPQSMKDYIATPKPNGYQSLHTKVLPFLYENTFRLEIQIRSEDMD 182
ICYHVLGL H++W P+P+++KDYIATPKPNGYQSLHT IR+E+MD
Sbjct: 239 ICYHVLGLVHEIWKPIPRTVKDYIATPKPNGYQSLHTT---------------IRTEEMD 283

Query: 183 IIAERGIAAHYSGRTVCTSLSTEA----GSHKGSSSFMNNVDLARRVSWLNAIREWQEEF 350
+IAERGIA +Y+G+++ T L A + +G + +NN D A RV WLNAIREWQEEF
Sbjct: 284 LIAERGIAVYYNGKSLSTGLVGNAVPLGRNSRGKTGCLNNADFALRVGWLNAIREWQEEF 343

Query: 351 VGNMSAQEFVDTVTGDLLGSRVFVFTPKGEIKNLPKGATVIDYAYQIHTDLGNNMVAAKV 530
VGNMS++EFVDT+T DLLGSRVFVFTPKGEIKNLPKGATV+DYAY IHT++GN MVAAKV
Sbjct: 344 VGNMSSREFVDTITRDLLGSRVFVFTPKGEIKNLPKGATVVDYAYLIHTEIGNKMVAAKV 403

Query: 531 NGNLVSPTHTLANAEVVEVITYNGLSIKSVFHRHRQWLQHAKT 659
NGNLVSPTH L NAEVVE++TYN LS KS F RH+QWLQHAKT
Sbjct: 404 NGNLVSPTHVLENAEVVEIVTYNALSSKSAFQRHKQWLQHAKT 446